.: Selected Publications:.

(U denotes undergraduate researcher mentored)

Boyce. C.K., Ibarra, D.E., Nelsen, M.P., and D’Antonio, M.P. 2023. Nitrogen-based symbioses, phosphorus availability, and accounting or a modern world more productive than the Paleozoic. Geobiology 21: 86-101.
Data & CodeGitHub

Nelsen, M.P., Lücking, R., Boyce, C.K., Lumbsch, H.T. and R.H. Ree. 2020. The macroevolutionary dynamics of symbiotic and phenotypic diversification in lichens. Proceedings of the National Academy of Sciences, USA 117: 21495-21503.
Media CoveragePress
Data & CodeGitHub

Nelsen, M.P., Lücking, R., Boyce, C.K., Lumbsch, H.T. and R.H. Ree. 2020. No support for the emergence of lichens prior to the evolution of vascular plants. Geobiology 18: 3-13.
Media CoveragePress | Press | Press | Press | Press | Press |
Data & CodeGitHub

Nelsen, M.P., Ree, R. and C.S. Moreau. 2018. Ant-plant interactions evolved through increasing interdependence. Proceedings of the National Academy of Sciences, USA 115: 12253-12258. 
Media CoveragePress | Press | Press | Press |
Data & CodeDryad | GitHub |

Nelsen, M.P., DiMichele, W.A., Peters, S.E. and C.K. Boyce. 2016. Delayed fungal evolution did not cause the Paleozoic peak in coal production. Proceedings of the National Academy of Sciences, USA 113: 2442-2447. 
Media CoveragePress | Press | Press | Press | Press |

.: Full Publication List :.

(U denotes undergraduate researcher mentored)


Meyer, A.R., Valentin, M., Liulevicius, L., McDonald, T., Nelsen, M.P., Pengra, J., Smith, R.J., and Stanton, D. 2023. Climate warming causes photobiont degradation and C starvation in a boreal sentinal lichen. American Journal of Botany, in press.

Boyce. C.K., Ibarra, D.E., Nelsen, M.P., and D’Antonio, M.P. 2023. Nitrogen-based symbioses, phosphorus availability, and accounting or a modern world more productive than the Paleozoic. Geobiology 23: 86-101.
Data & CodeGitHub


Phraphuchamnong, P., Nelsen, M.P., Distefano, I., Mercado-Diaz, J., Parnmen, S., Rangsiruji, A., Buaruang, K., Lücking, R., and Lumbsch, H.T. 2022. A new species of Megalaria (Ramalinaceae, Ascomycota) from Thailand, and recognition of subgenus Catillochroma. MycoKeys 93: 149-163.

Nelsen, M.P. and Boyce. C.K. 2022. What to do with Prototaxites? International Journal of Plant Sciences 183: 556-565.

Nelsen, M.P., Leavitt, S.D., UHeller, K., Muggia, L., and Lumbsch. H.T. 2022. Contrasting patterns of climatic niche divergence in Trebouxia—a clade of lichen-forming algae. Frontiers in Microbiology 13: 791546.
Media CoveragePress | Press | Press |
Data & CodeGitHub


Nelsen, M.P., Leavitt, S.D., UHeller, K., Muggia, L., and Lumbsch. H.T. 2021. Macroecological diversification and convergence in a clade of keystone symbionts. FEMS Microbiology Ecology 97: fiab072.
Data & CodeGitHub

Nelsen, M.P. 2021. Sharing and double-dating in the lichen world. Molecular Ecology 30: 1751-1754.


Nelsen, M.P., Lücking, R., Boyce, C.K., Lumbsch, H.T. and R.H. Ree. 2020. The macroevolutionary dynamics of symbiotic and phenotypic diversification in lichens. Proceedings of the National Academy of Sciences, USA 117: 21495-21503.
Media CoveragePress
Data & CodeGitHub

Nelsen, M.P., Lücking, R., Boyce, C.K., Lumbsch, H.T. and R.H. Ree. 2020. No support for the emergence of lichens prior to the evolution of vascular plants. Geobiology 18: 3-13.
Media CoveragePress | Press | Press | Press | Press | Press |
Data & CodeGitHub

Nelsen, M.P.  and Lumbsch, H.T. 2020. A data-driven evaluation of lichen climate change indicators in Central Europe. Biodiversity and Conservation 29: 3959-3971.
Data & CodeGitHub

Lindgren, H., Moncada, B., Lücking, R., Magain, N., Simon, A., Goffinet, B., Sérusiaux, E., Nelsen, M.P., Mercado-Díaz, J.A., Widhelm, T.J., and Lumbsch, H.T. 2020. Cophylogenetic patterns in algal symbionts correlate with repeated symbiont switches during diversification and geographic expansion of lichen-forming fungi in the genus Sticta (Ascomycota, Peltigeraceae). Molecular Phylogenetics and Evolution 150: 106860.

Muggia, L., Nelsen, M.P., Kirika, P.M., Barreno, E., Beck, A., Lindgren, H., Lumbsch, H.T., Leavitt, S. D., and Trebouxia working group. 2020. Formally described species woefully underrepresent phylogenetic diversity in the common lichen photobiont genus Trebouxia (Trebouxiophyceae, Chlorophyta): An impetus for developing an integrated taxonomy. Molecular Phylogenetics and Evolution 149: 106821.

Põlme, S., Abarenkov, K., Nilsson, R.H., Lindahl, B.D., Clemmensen, K., Kauserud, H., Nguyen, N., Kjoller, R., Bates, S.T., Baldrian, P.,Frøslev, T.G., Adojaan, K., Vizzini, A., Suija, A., Pfister, D., Baral, H.-O., Järv, H., Madrid, H., Nordén, J., Liu, J.-K., Pawlowska, J.,Põldmaa, K., Pärtel, K., Runnel, K., Hansen, K., Larsson, K.-H., Hyde, K.D., Sandoval-Denis, M., Smith, M.E., Toome-Heller, M.,Wijayawardene, N.N., Menolli Jr., N., Reynolds, N.K., Drenkhan, R., Maharachchikumbura, S.S.N., Gibertoni, T.B., Læssøe, T., Davis, W.,Tokarev, Y., Corrales, A., Soares, A.M. Agan, A., Machado, A.R., Argüelles-Moyao, A., Detheridge, A., de Meiras-Ottoni, A., Verbeken,A., Dutta, A.K., Cui, B.-K., Pradeep, C.K., Marín, C., Stanton, D., Gohar, D., Wanasinghe, D.N., Otsing, E., Aslani, F., Griffith, G.W.,Lumbsch, T.H., Grossart, H.-P., Masigol, H., Timling, I. Hiiesalu, I., Oja, J., Kupagme, J.Y., Geml, J., Alvarez-Majarrez, J., Ilves, K., Loit,K., Adamson, K., Nara, K. Küngas, K., Rojas-Jimenez, K., Bitenieks, K., Irinyi, L., Nagy, L.L., Soonvald, L., Zhou, L.-W., Wagner, L., Aime,M.C., Öpik, M., Mujica, M.I., Metsoja, M., Ryberg, M., Vasar, M., Murata, M., Nelsen, M.P., Cleary, M., Samarakoon, M.C., Doilom,M., Bahram, M., Hagh- Doust, N., Dulya, O., Johnston, P., Kohout, P., Chen, Q., Tian, Q., Nandi, R., Amiri, R., Perera, R.H., dos SantosChikowski, R., Mendes-Alvarenga, R.L., Garibay-Orijel, R., Gielen, R., Phookamsak, R., Jayawardena, R.S., Rahimlou, S., Karunarathna, S.C., Tibpromma, S., Brown, S.P., Sepp, S.-K., Mundra, S., Luo, Z.-H., Bose, T., Vahter, T., Netherway, T., Yang, T., May, T., Varga, T., Li,W., Coimbra, V.R.M., de Oliveira, V.R.T., de Lima, V.X., Mikryukov, V.S., Lu, Y. Matsuda, Y., Miyamoto, Y., Kõljalg, U. and Tedersoo, L. 2020. FungalTraits: a user-friendly traits database of fungi and fungus-like stramenopiles. Fungal Diversity 105: 1-16.


Huang, J.-P., Kraichak, E., Leavitt, S.D., Nelsen, M.P.  and H.T. Lumbsch. 2019. Accelerated diversification in three diverse families of morphologically complex lichen-forming fungi links to major geohistorical events. Scientific Reports 9: 8518.
Media CoveragePress | Press | Press | Press |


Nelsen, M.P., Ree, R. and C.S. Moreau. 2018. Ant-plant interactions evolved through increasing interdependence. Proceedings of the National Academy of Sciences, USA 115: 12253-12258.
Media CoveragePress | Press | Press | Press |
Data & CodeDryad | GitHub |

Kraichack, E., Huang, J.-P., Nelsen, M., Leavitt, S.D. and H.T. Lumbsch. 2018. A revised classification of orders and families in the two major subclasses of Lecanoromycetes (Ascomycota) based on a temporal approach. Botanical Journal of the Linnean Society 188: 233-249.

Will-Wolf, S., Jovan, S., Nelsen, M.P., Trest, M.T., Rolih, K.M. and A.H. Reis. 2018. Lichen-indices assess local climate and air quality status in the Mid-Atlantic region, USA. Bryologist 121: 461-479.

Lücking, R. & M.P. Nelsen. 2018. Ediacarans, protolichens and lichen-derived Penicillium: a critical reassessment of the evolution of lichenization in fungi. Pp. 551-590 in: Krings, M., Harber, C.J., Cuneo, N.R. and G.W. Rothwell (Eds.), Transformative paleobotany: papers to commemorate the life and legacy of Thomas N. Taylor. Academic Press, London, UK.

Leavitt, S.D., Westberg, M., Nelsen, M.P., Elix, J.A., Timdal, E., Sohrabi, M., St. Clair, L.L., Williams, L., Wedin, M. and H.T. Lumbsch. 2018. Multiple, distinct intercontinental lineages but isolation of Australian populations in a cosmopolitan lichen-forming fungal taxon, Psora decipiens (Psoraceae, Ascomycota). Frontiers in Microbiology 9: 283.


Nelsen, M.P., Lücking, R., Cáceres, M.E.S., Aptroot, A. and H.T. Lumbsch. 2017. Assessing the phylogenetic placement and redundancy of Aspidotheliaceae (Ascomycota), an orphaned family of lichen-forming fungi. Systematics and Biodiversity 15: 63-73.


Nelsen, M.P., DiMichele, W.A., Peters, S.E. and Boyce, C.K. 2016. Delayed fungal evolution did not cause the Paleozoic peak in coal production. Proceedings of the National Academy of Sciences, USA 113: 2442-2447. 
Media CoveragePress | Press | Press | Press | Press |

Sanders, W.B., Pérez-Ortega, S., Nelsen, M.P., Lücking, R. and A. de Los Ríos. 2016. Heveochlorella (Trebouxiophyceae): a little-known genus of unicellular green algae outside the Trebouxiales emerges unexpectedly as a major clade of lichen photobionts in foliicolous communities. Journal of Phycology 52: 840-853.

Leavitt, S.D., Kraichak, E., Vondrak, J., Nelsen, M.P., Sohrabi, M., Pérez-Ortega, S., St. Clair, L.L. and H.T. Lumbsch. 2016. Cryptic diversity and symbiont interactions in rock-posy lichens. Molecular Phylogenetics and Evolution 99: 261-274.

Lücking, R., Nelsen, M.P., Aptroot, A., Barillas de Klee, R., Bawingan, P.A., Benattie, M.N., Binh, N.Q., Bungartz, F., Cáceres, M.E.S., da Silva Canêz, L., Chaves, J.L., Ertz, D., Esquivel, R.E., Ferraro, L.I., Grijalva, A., Gueidan, C. Hernández, J.E., Knight, A., Lumbsch, H.T., Marcelli, M.P., Mercado-Díaz, J.A., Moncada, B., Morales, E.A., Naksuwankul, K., Orozco, T., Parnmen, S., Rivas Plata, E., Salazar-Allen, N., Spielmann, A.A. and N. Ventura. 2016. A phylogenetic framework for reassessing generic concepts and species delimitation in the lichenized family Trypetheliaceae (Ascomycota: Dothideomycetes). Lichenologist 48: 739-762.

Lücking, R., Nelsen, M.P., Aptroot, A., Benatti, M.N., Binh, N.Q., Gueidan, C., Gutiérrez, M.C., Jungbluth, P., Lumbsch, H.T., Marcelli, M.P., Moncada, B., Naksuwankul, K., Orozco, T., Salazar-Allen, N. and D. Upreti. 2016. A pot-pourri of new species of Trypetheliaceae resulting from molecular phylogenetic studies. Lichenologist 48: 639-660.

Mark, K., Saag, L., Leavitt, S.D., Will-Wolf, S., Nelsen, M.P., Tõrra, T., Randlane, T. and H.T. Lumbsch. 2016. Evaluation of traditionally circumscribed species in the lichen-forming genus Usnea, section Usnea(Parmeliaceae, Ascomycota) using a six locus dataset. Organisms, Diversity & Evolution 16: 497-524.

Luangsuphabool, T., Piapukiew, J., Parnmen, S., Nelsen, M.P., Lumbsch, H.T. and Sangvichien, E. 2016. Diversity of the Trypethelium eluteriae group in Thailand (Ascomycota, Trypetheliales). Lichenologist 48: 53-60.


Kraichak, E., Divakar, P.K., Crespo, A., Leavitt, S.D., Nelsen, M.P., Lücking, R. and Lumbsch, H.T. 2015. A tale of two hyper-diversities: diversification dynamics of the two largest families of lichenized fungi. Scientific Reports 5: 10028. 

Leavitt, S.D., Kraichak, E., Nelsen, M.P., Altermann, S., Divakar, P.K., Alors, D., Esslinger, T.L., Crespo, A. and Lumbsch, H.T. 2015. Fungal specificity and selectivity for algae play a major role in determining lichen partnerships across diverse ecogeoraphic regions in the lichen-forming family Parmeliaceae (Ascomycota). Molecular Ecology 24: 3779-3797.

Divakar, P.K., Crespo, A., Wedin, M., Leavitt, S.D., Myllys, L., McCune, B., Randlane, T., Bjerke, J.W., Ohmura, Y., Schmitt, I., Boluda, C.G., Alors, D., Roca-Valiente, B., Del-Prado, R., Ruibal, C., Buaruang, K., Núñez-Zapata, J., Amo de Paz, G., Rico, V.J., Molina, M.C., Elix, J.A., Esslinger, T.L., Tronstad, I.K.K., Lindgren, H., Ertz, D., Gueidan, C., Saag, L., Tõrra, T., Singh, G., Dal Grande, F., Parnmen, S., Beck, A., Navarro Benatti, M., Blanchon, D., Candan, M., Clerc, P., Goward, T., Grube, M., Hodkinson, B.P., Hur, J.-S., Kantvilas, G., Kirika, P.M., Lendemer, J., Mattsson, J.-E., Messuti, M.I., Miadlikowska, J., Nelsen, M., Ohlson, J.I., Pérez-Ortega, S., Saag, A., Sipman, H.J.M., Sohrabi, M., Thell, A., Thor, G., Truong, C., Yahr, R., Upreti, D.K., Hawksworth, D.L., Cubas, P. and Lumbsch, H.T. Evolution of complex symbiotic relationships in a morphologically derived family of lichen-forming fungi. New Phytologist 208: 1217-1226. 

Will-Wolf, S., Makholm, M.M., Nelsen, M.P., Trest, M.T., Reis, A.H. and Jovan, S. 2015. Element analysis of two common macrolichens supports bioindication of air pollution and lichen response in rural midwestern U.S.A. Bryologist 118: 371-384.

Kraichak, E., Lücking, R. Aptroot, A., Beck, A., Dornes, P., John, V., Lendemer, J.C., Nelsen, M.P., Neuwirth, G., Nutakki, A., Parnmen, S., Sohrabi, M., Tønsberg, T. and Lumbsch, H.T. 2015. Hidden diversity in the morphologically variable script lichen (Graphis scripta) complex (Ascomycota, Ostropales, Graphidaceae). Organisms, Diversity & Evolution 15: 447-458. 


Dal Grande, F., Beck, A., Cornejo, C., Singh, G., Cheenacharoen, S., Nelsen, M.P. and Scheidegger, C. 2014. Molecular phylogeny and symbiotic selectivity of the green algal genus Dictyochloropsis s.l. (Trebouxiophyceae): a polyphyletic and widespread group forming photobiont-mediated guilds in the lichen family Lobariaceae. New Phytologist 202: 455-470. 

Altermann, S., Leavitt, S.D., Goward, T., Nelsen, M.P. and Lumbsch, H.T. 2014. How do you solve a problem like Letharia? A new look at cryptic species in lichen-forming fungi using bayesian clustering and SNPs from multilocus sequence data. PLoS ONE 9: e97556. 

Nelsen, M.P., Lücking, R., Aptroot, A., Andrew, C.J., Cáceres, M.E.S., Rivas Plata, E., Gueidan C., da Silva Canêz, L., Knight, A., Ludwig, L.R., Mercado-Díaz, J., Parnmen, S. and Lumbsch, H.T. 2014. Elucidating phylogenetic relationships and genus-level classification within the fungal family Trypetheliaceae (Dothideomycetes: Ascomycota). Taxon 63: 974-992. 

Nelsen, M.P., Lücking, R., Andrew, C.J., Aptroot, A., Cáceres, M.E.S., Mercado-Díaz, J. and Lumbsch, H.T. 2014. Molecular phylogeny reveals the true colors of Myeloconidaceae (Ascomycota: Ostropales). Australian Journal of Systematic Botany 27: 38-47. 

Wijayawardene, N.N., Crous, P.W., Kirk, P.M., Hawksworth, D.L., Boonmee, S., Braun, U., Dai, D.-Q., D’souza, M.J., Diederich, P., Dissanayake, A., Doilom, M., Hongsanan, S., Jones, E.B.G., Groenewald, J.Z., Jayawardena, R., Lawrey, J.D., Liu, J.-K., Lücking, R., Madrid, H., Manamgoda, D.S., Muggia, L., Nelsen, M.P., Phookamsak, R., Suetrong, S., Tanaka, K., Thambugala, K.M., Wanasinghe, D.N., Wikee, S., Zhang, Y., Aptroot, A., Ariyawansa, H.A., Bahkali, A.H., Bhat, D.J., Gueidan, C., Chomnunti, P., De Hoog, G.S., Knudsen, K., Li, W.-J., McKenzie, E.H.C., Miller, A.N., Phillips, A.J.L., Piątek, M., Raja, H.A., Shivas, R.S., Slippers, B., Taylor, J.E., Tian, Q., Wang, Y., Woudenberg, J.H.C., Cai, L., Jaklitsch, W. and Hyde, K.D. 2014. Naming and outline of Dothideomycetes-2014 including proposals for the protection or suppression of generic names. Fungal Diversity 69 1-55.


Hyde, K.D., Liu, J.-K., Binder, M., Aryawansha, H., Boehm, E., Boonmee, S., Braun, U., Chomnunti, P., Crous, P.W., Dai, D., Diederich, P., Dissanayake, A., Doilom, M., Doveri, F., Hongsanan, S., Jayawardena, R., Lawrey, J.D., Li, Y.M., Liu, K., Lücking, R., Monkai, J., Nelsen, M.P., Phookamsak, R., Muggia, L., Pang, K.-L., Senanayake, I., Shearer, C.A., Wijayawardene, N., Wu, X.H., Thambugala, K.M., Hongsanan, S., Suetrong, S., Tanaka, K., Wikee, S., Yaharoen, S., Zhang, Y., Aguirre-Hudson, B., Alias, S.A., Aptroot, A., Bezerra, J.L., Bhat, J.D., Camporesi, E., Chukeatirote, E., Dakshitha, G., De Hoog, S., Gueidan, C., Hawksworth, D.L., Hirayama, K., Kang, J.C., Knudsen, K., Li, W.J., Liu, Z., McKenzie, E.H.C., Miller, A., Philipps, A., Raja, H.A., Qing, T., Taylor, J., Wang, Y., Yan, J., Zhang, M. and Jones, E.B.G. 2013. Families of Dothideomycetes. Fungal Diversity 63: 1-313.    

Lücking, R., Breuss, O., Nelsen, M.P., Navarro, E., Chaves, J.L., Trest, M. and Will-Wolf, S. 2013. Porina squamulifera (lichenized Ascomycota: Porinaceae), a new species from tropical rainforest in Costa Rica with unique thallus morphology. Herzogia 26: 223-230. 

Cáceres, M.E.S., Aptroot, A., Nelsen, M.P. and Lücking, R. 2013. Pyrenula sanguinea (lichenized Ascomycota, Pyrenulaceae), a new species with unique, trypethelioid ascomata and complex pigment chemistry. Bryologist 116: 350-357. 

Nelsen, M.P., Thell, A., Leavitt, S.D., Hampton-Miller, C.J. and Lumbsch, H.T. 2013. Molecular data motivate a reappraisal of Masonhalea (Parmeliaceae, Lecanorales). Lichenologist 45: 729-738. 

Lücking, R., dal Forno, M., Lawrey, J.D., Hernández, J.E., Marcelli, M.P., Moncada, B., Morales, E., Nelsen, M.P., Paz, E., Salcedo, L., Spielmann, A., Wilk, K., Will-Wolf, S. and Yánez-Ayabaca, A. 2013. Nine new species of lichenized Basidiomycota in the genera Dictyonema and Cora (Agaricales: Hygrophoraceae). Phytotaxa 139: 1-38.

Aptroot, A.,Nelsen, M.P. and S. Parnmen. 2013. Marcelaria, a new genus for the Laurera purpurina group in the Trypetheliaceae (Ascomycota: Dothideomycetes). Glalia 5: 1-14.

Leavitt, S.D., Nelsen, M.P., Lumbsch, H.T., Johnson, L.A. and St. Clair, L.L. 2013b. Symbiont flexibility in subalpine rock shield lichen communities in the Southwestern USA. Bryologist 116: 149-161.

Leavitt, S.D., Esslinger, T.L., Nelsen, M.P. and Lumbsch, H.T. 2013a. Further species diversity in Neotropical Oropogon (Lecanoromycetes: Parmeliaceae) in Central America. Lichenologist 45: 553-564.

Rivas Plata, E., Parnmen, S., Staiger, B., Mangold, A., Frisch, A., Weerakoon, G., Hernández, J.E., Cáceres, M.E.S., Kalb, K., Sipman, H.J.M., Common, R.S., Nelsen, M.P., Lücking, R. and Lumbsch, H.T. 2013. A molecular phylogeny of Graphidaceae (Ascomycota, Lecanoromycetes, Ostropales) including 428 species. MycoKeys 6: 55-94.


Nelsen, M.P., Lücking, R., Andrew, C.J., Rivas Plata, E., Chaves, J.L., Cáceres, M.E.S. and Ventura, N. 2012. Dismantling Herpothallon: Herpothallon antillarum (Arthoniomycetes: Arthoniaceae) is a member of the genus Diorygma (Lecanoromycetes: Graphidaceae). Bryologist 115: 313-321.

Lawrey, J., Diederich, P., Nelsen, M.P., Freebury, C., Van den Broeck, D., Sikaroodi, M. and Ertz, D. 2012. Phylogenetic placement of lichenicolous Phoma species in the Phaeosphaeriaceae (Pleosporales, Dothideomycetes). Fungal Diversity 55: 195-213.

Sweetwood, G., Lücking, R., Nelsen, M.P. and Aptroot, A. 2012. Ascospore ontogeny in megalosporous Trypetheliaceae and Graphidaceae (Ascomycota: Dothideomycetes and Lecanoromycetes) suggests phylogenetic relationships and ecological constraints. Lichenologist 44: 277-296.


Nelsen, M.P., Rivas Plata, E., Andrew, C.J., Lücking, R. and Lumbsch, H.T. 2011. Phylogenetic diversity of trentepohlialean algae associated with lichen-forming fungi. Journal of Phycology 47: 282-290.

Thüs, H., Muggia, L., Pérez-Ortega, S., Favero-Longo, S.E., Joneson, S., O’Brien, H., Nelsen, M.P., Duque-Thüs, R., Grube, M., Friedl, T., Brodie, J., Andrew, C.J., Lücking, R., Lutzoni, F. and Gueidan, C. 2011. Revisiting photobiont diversity in the lichen family Verrucariaceae (Ascomycota). European Journal of Phycology 46: 399-415.

Will-Wolf, S., Nelsen, M.P. and M.T. Trest. 2011. Does morphological response of four common lichen species to pollution, shade and landscape pattern predict long-term changes in distribution. Bibliotheca Lichenologica 106: 375-386.

Nelsen, M.P., Lücking, R., UMbatchou, J.S., Andrew, C.J., Spielmann, A.A. and Lumbsch, H.T. 2011. New insights into relationships of lichen-forming Dothideomycetes. Fungal Diversity 51: 155-162.

Nelsen, M.P., UChavez, N., Sackett-Hermann, E., Thell, A., Randlane, T., Divakar, P.K., Rico, V.J. and Lumbsch, H.T. 2011. The cetrarioid core group revisited (Lecanorales: Parmeliaceae). Lichenologist 43: 537-551.

Lücking, R., Seavey, F., Common, R.S., Beeching, S.Q., Breuss, O., Buck, W.R., Crane, L., Hodges, M., Hodkinson, B.P., Lay, E., Lendemer, J.C., McMullin, R.T., Mercado Díaz, J.A., Nelsen, M.P., Rivas Plata, E., Safranek, W., Sanders, W.B., Schaefer Jr., H.P. and Seavey, J. 2011. The lichens of Fakahatchee Strand Preserve State Park, Florida: proceedings from the 18th Tuckerman workshop. Bulletin of the Florida Museum of Natural History 49: 127-186.
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Lumbsch, H.T., Ahti, T., Altermann, S., Amo De Paz, G., Aptroot, A., Arup, U., Bárcenas Peña, A., Bawingan, P.A., Benatti, M.N., Betancourt, L., Björk, C.R., Boonpragob, K., Brand, M., Bungartz, F., Cáceres, M.E.S., Candan, M., Chaves, J.L., Clerc, P., Common, R., Coppins, B.J., Crespo, A., Dal-Forno, M., Divakar, P.K., Duya, M.V., Elix, J.A., Elvebakk, A., Fankhauser, J.D., Farkas, E., Itatí Ferraro, L., Fischer, E., Galloway, D.J., Gaya, E., Giralt, M., Goward, T., Grube, M., Hafellner, J., Hernández M., J.E., De Los Angeles Herrera Campos, M., Kalb, K., Kärnefelt, I., Kantvilas, G., Killmann, D., Kirika, P., Knudsen, K., Komposch, H., Kondratyuk, S., Lawrey, J.D., Mangold, A., Marcelli, M.P., McCune, B., Ines Messuti, M., Michlig, A., Miranda González, R., Moncada, B., Naikatini, A., Nelsen, M.P., Øvstedal, D.O., Palice, Z., Papong, K., Parnmen, S., Pérez-Ortega, S., Printzen, C., Rico, V.J., Rivas Plata, E., Robayo, J., Rosabal, D., Ruprecht, U., Salazar Allen, N., Sancho, L., Santos De Jesus, L., Santos Vieira, T., Schultz, M., Seaward, M.R.D., Sérusiaux, E., Schmitt, I., Sipman, H.J.M., Sohrabi, M., Søchting, U., Zeuthen Søgaard, M., Sparrius, L.B., Spielmann, A., Spribille, T., Sutjaritturakan, J., Thammathaworn, A., Thell, A., Thor, G., Thüs, H., Timdal, E., Truong, C., Türk, R., Umaña Tenorio, L., Upreti, D.K., Van Den Boom, P., Vivas Rebuelta, M., Wedin, M., Will-Wolf, S., Wirth, V., Wirtz, N., Yahr, R., Yeshitela, K., Ziemmeck, F., Wheeler, T. and Lücking, R. 2011. One hundred new species of lichenized fungi: a signature of undiscovered global diversity. Phytotaxa 18: 1-127.
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Lawrey, J.D., Diederich, P, Nelsen, M.P., Sikaroodi, M., Gillevet, P.M., Brand, M. and Van den Boom, P. 2011. The obligately lichenicolous genus Lichenoconium represents a novel lineage in the Dothideomycetes. Fungal Biology 115: 176-187.


Nelsen, M.P., Lücking, R. Rivas Plata, E. and UMbatchou, J.S. 2010. Heiomasia, a new genus in the lichen-forming family Graphidaceae (Ascomycota: Lecanoromycetes: Ostropales) with disjunct distribution in Southeastern North America and Southeast Asia. Bryologist 113: 742-751.

Will-Wolf, S., Nelsen, M.P. and Trest, M.T. 2010. Responses of small foliose lichen species to landscape pattern, light regime, and air pollution from a long-term study in upper Midwest USA. Bibliotheca Lichenologica 105: 167-182.


Nelsen, M.P. and Gargas, A. 2009. Symbiont flexibility in the lichen Thamnolia vermicularis (Pertusariales: Icmadophilaceae). Bryologist 112: 404-417.

Nelsen, M.P. and Gargas, A. 2009. Assessing clonality and chemotype monophyly in Thamnolia (Pertusariales: Icmadophilaceae). Bryologist 112: 42-53

Nelsen, M.P., Lücking, R., Grube, M., UMbatchou, J.S., Muggia, L., Rivas Plata, E. and Lumbsch, H.T. 2009. Unravelling the phylogenetic relationships of lichenised fungi in Dothideomyceta. Studies in Mycology 64: 135-144.

Schoch, C.L., Crous, P.W., Groenewald, J.Z., Boehm, E.W.A., Burgess, T.I., de Gruyter, J., de Hoog, G.S., Dixon, L.J., Grube, M., Gueidan, C., Harada, Y., Hatakeyama, S., Hirayama, K., Hosoya, T., Huhndorf, S.M., Hyde, K.D. Jones, E.B.G., Kohlmeyer, J., Kruys, Å., Li, Y.M., Lücking, R. Lumbsch, H.T., Marvanová, L., UMbatchou, J.S., McVay, A.H., Miller, A.N., Mugambi, G.K., Muggia, L., Nelsen, M.P., Nelson, P., Owensby, C.A., Phillips, A.J.L., Phongpaichit, S., Pointing, S.B., Pujade-Renaud, V., Raja, H.A., Rivas Plata, E., Robbertse, B., Ruibal, C., Sakayaroj, J., Sano, T., Selbmann, L., Shearer, C.A., Shirouzu, T., Slippers, B., Suetrong, S., Tanaka, K., Volkmann-Kohlmeyer, B., Wingfield, M.J., Wood, A.R., Woudenberg, J.H.C., Yonezawa, H., Zhang, Y. and Spatafora, J.W. 2009. A class-wide phylogenetic assessment of Dothideomycetes. Studies in Mycology 64: 1-15.

Payyavula, R.S., Babst, B.A., Nelsen, M.P., Harding, S.A. and Tsai, C.-J. 2009. Glycosylation-mediated phenylpropanoid partitioning in Populus tremuloides cell cultures. BMC Plant Biology 9: 151.

Schmitt, I., Crespo, A., Divakar, P.K., Fankhauser, J., Hermann-Sackett, E., Kalb, K., Nelsen, M.P., Nelson, N.A., Rivas-Plata, E., Shimp, A.D., Widhelm, T. and Lumbsch, H.T. 2009. New primers for promising single-copy genes in fungal phylogenetics and systematics. Persoonia 23: 35-40.

Zielinska, B., Killian, M., Kadmiel, M., Nelsen, M. and Haut Donahue, T.L. 2009. Meniscal tissue explants response depends on level of dynamic compressive strain. Osteoarthritis and Cartilage 17: 754-760.

Cseke, L.J., Tsai, C.-J., Rogers, A., Nelsen, M.P., White, H.L., Karnosky, D.F. and Podila, G.K. 2009. Transcriptomic comparison in the leaves of two aspen genotypes having similar carbon assimilation rates but different partitioning patterns under elevated [CO2]. New Phytologist 182: 891-911.


Nelsen, M.P. and Gargas, A. 2008. Dissociation and horizontal transmission of co-dispersing lichen symbionts in the genus Lepraria (Lecanorales: Stereocaulaceae). New Phytologist 177: 264-275.

Will-Wolf, S. and Nelsen, M.P. 2008. How have Wisconsin’s lichen communities changed?  Pp. 135-150 in: Waller, D.M. and Rooney, T.P. (Eds.), The vanishing present: Ecological change in Wisconsin. University of Chicago Press, Chicago, USA.

Nelsen, M.P. and Gargas, A. 2008. Phylogenetic distribution and evolution of secondary metabolites in the lichenized fungal genus Lepraria (Lecanorales: Stereocaulaceae). Nova Hedwigia 86:115-131.

Donahue, S.W., McGee, M.E., Bradford, R.M., Tsai, C.-J., Nelsen, M.P. and Simoni, K. 2008. American black bear parathyroid hormone as therapy for osteoporosis. Pp. 25-36 in: Lovegrove, B.G. and McKechnie, A.E. (Eds.), Hypometabolism in animals: hibernation, torpor and cryobiology. University of Kwa-Zulu Natal, Pietermaritzburg, South Africa. 

Nelsen, M.P., Lumbsch, H.T., Lücking, R. and Elix, J.A. 2008. Further evidence for the polyphyly of Lepraria (Lecanorales: Stereocaulaceae). Nova Hedwigia 87: 361-371.

Lumbsch, H.T., Nelsen, M.P. and Lücking, R. 2008. The phylogenetic position of Haematommataceae (Lecanorales, Ascomycota) with notes on secondary chemistry and species delimitation. Nova Hedwigia 86: 105-114.


Nelsen, M.P., Will-Wolf, S. and Gargas, A. 2007. One-hundred years of change in the corticolous macrolichens of Madison, Wisconsin. Evansia 24: 108-112.

Nelsen, M.P., Lücking, R., Umaña, L., Trest, M.T., Will-Wolf, S., Chaves, J.L., and Gargas, A. 2007. Multiclavula ichthyiformis (Fungi: Basidiomycota: Cantharellales: Clavulinaceae), a remarkable new basidiolichen from Costa Rica. American Journal of Botany 94: 1289-1296.

Nelsen, M.P. 2007. Noteworthy collections in Michigan: Multiclavula vernalis (Schw.) Petersen and Multiclavula mucida (Fr.) Petersen. Michigan Botanist 46: 124-126.


Nelsen, M.P. and Gargas, A. 2006. Actin type I introns offer potential for increasing phylogenetic resolution in Asterochloris (Chlorophyta: Trebouxiophyceae). Lichenologist 38: 435-440.

Nelsen, M.P., Lücking, R., Chaves, J. L., Sipman, H. J. M., Umaña, L. and Navarro, E. 2006. A first assessment of the Ticolichen biodiversity inventory in Costa Rica: the genus Haematomma (Lecanorales: Lecanoraceae). Lichenologist 38: 251-262.

Nelsen, M.P. 2006. An addition to the lichen flora of New Jersey: the basidiolichen Multiclavula vernalis (Schw.) Petersen. Evansia 23: 11.

Lücking, R, Aptroot, A., Umaña, L., Chaves, J. L., Sipman, H. J. M. and Nelsen, M.P. 2006. A first assessment of the Ticolichen biodiversity inventory in Costa Rica: The genus Gyalideopsis (Ostropales: Gomphillaceae). Lichenologist 38: 131-160.


Nelsen, M.P. 2005. The basidiolichen Multiclavula mucida (Fr.) Petersen: new to Michigan. Michigan Botanist 44: 192-193.

Nelsen, M.P. 2005. Additions to the lichen flora of Wisconsin with new records of rare species. Michigan Botanist 44: 188-191.

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